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- Estimation of stand crown cover using a generalized crown diameter model: application for the analysis of portuguese cork oak stands stocking evolutionPublication . Paulo, Joana Amaral; Faias, Sónia; Ventura-Giroux, Célia; Tomé, MargaridaA generalized non-linear tree crown diameter model was developed with the aim of allowing the computation of tree crown diameter over a large range of tree dimensions, and allowing its inclusion in forest growth and yield models. The model was formulated to provide biological meaning to the predicted values. Due to the nested structure of the data analyzed (trees within stands), both mixed- and fixed-effect models were developed. Since tree crown diameter is not frequently measured in forest inventories, the validation of the mixed model was carried out by considering the population specific response. The results demonstrate that when the measurements required for the mixed model calibration are not available, the use of the fixed effect model results in less biased and more accurate estimates. The fixed model was applied to the data from the two last Portuguese National Forest Inventories (NFI) to analyze the change in stand crown cover and assess the stocking evolution of cork oak stands in Portugal between 1996 and 2006. Results showed an increase in the frequency of stands with crown cover lower than 20%, as well as a decrease in the frequency of stands with crown cover between 20 and 40%. Average crown cover values were significantly different in the two NFI, with a decrease from 28.0 to 26.5% over the considered period
- Predicting site index from climate and soil variables for cork oak (Quercus suber L.) stands in PortugalPublication . Paulo, Joana Amaral; Palma, J.H.N.; Gomes, Alberto Azevedo; Faias, Sónia; Tomé, José; Tomé, MargaridaSite productivity, assessed through site index, was modelled using partial least squares regression as a function of soil and climatic variables. Two alternative models were developed: a full model, considering all available explanatory variables, and a reduced model, considering only variables that can be obtained without digging a soil pit. The reduced model was used for mapping the site index distribution in Portugal, on the basis of existing digital cartography available for the whole country. The developed models indicate the importance of water availability and soil water holding capacity for site index value distribution. Site index was related to climate, namely evaporation and frost, and soil characteristics such as lithology, soil texture, soil depth, thickness of the A horizon and soil classification. The variability of the estimated values within the map (9.5–16.8 m with an average value of 13.4 m) reflects the impact of soil characteristics on the site productivity estimation. These variables should be taken into consideration during the establishment of new plantations of cork oak, and management of existing plantations. Results confirm the potential distribution of cork oak in coastal regions. They also suggest the existence of a considerable area, located both North and South of the Tagus river, where site indices values of medium (]13;15]) to high (]15;17]) productivity classes may be expected. The species is then expected to be able to have good productivity along the northern coastal areas of Portugal, where presently it is not a common species but where, according to historical records, it occurred until the middle of the sixteenth century. The present research focused on tree growth. Cork growth and cork quality distribution needs to be further researched through the establishment of long term experimental sites along the distribution area of cork oak, namely in the central and northern coastal areas of the country
- Understory effect on tree and cork growth in cork oak woodlandsPublication . Faias, Sónia; Paulo, Joana Amaral; Palma, J.H.N.; Tomé, MargaridaAim of study: Cork oak is one of the main forest tree species in Portugal that typically occurs in montado, where operational practices oriented to the tree, crop or animal management may influence several of the ecosystem components. This study aimed at contributing to fulfil the a lack of knowledge on the effect of these practices on the cork and wood growth, by comparing the wood diameter growth and the annual cork increment under two different understory management options. Material and methods: An experimental trial implemented on an uneven-aged cork oak pure stand during a cork rotation period of 9 years, was established with the specific goal of comparing understory management options: a yellow lupine pasture versus spontaneous vegetation. Cork samples were taken at the beginning and end of the period and were used to measure cork thickness and annual cork rings. The differences between treatments were assessed performing a non-parametric test and a more robust approach using linear mixed model. Precipitation and treatment levels were jointly considered on the analysis. Main results: A slight effect was found on the cork thickness regarding the treatment with lupine application. However, no distinct effect was found, regarding wood and the annual cork increment pattern. Additionally, annual cork ring width showed a positive correlation with precipitation and a negative correlation with ring age. Research highlights: The results of this study indicate no distinct pattern regarding the annual cork and wood increment when comparing the understory effect of yellow lupine pasture versus spontaneous vegetation
- Drivers for annual cork growth under two understory management alternatives on a podzolic cork oak standPublication . Faias, Sónia; Paulo, Joana Amaral; Firmino, Paulo N.; Tomé, MargaridaUnderstory management practices and stand density characteristics allow one to distinguish a cork oak traditional silvopastoral system (known as a montado) from a cork oak forest system. Although understanding the manner in which different management practices affect cork growth is imperative, there are still only a few outputs from experimental research that contribute to this knowledge. The effect of potential drivers on annual cork growth was analyzed using a linear mixed model approach. Two dimensions of drivers were considered: intraspecific competition, assessed by tree level distance-dependent indices; and interspecific competition, assessed by variables characterizing understory management. The present dataset was collected from an experimental trial established on a cork oak stand in Podzolic soil on the Tagus river basin, covering two different cork growth cycles over the period from 2003 to 2015. The adjusted models considered two understory management alternatives: spontaneous shrubs maintenance and forage application. In both models, annual precipitation displayed a positive effect on annual cork growth, as expected. However, no significant effect of intraspecific competition was found. Additionally, there was a positive effect on annual cork growth associated with the spontaneous shrubs growth and a negative effect associated with lupine presence; both effects linked to different cork ring ages’ thresholds. The study main contributions are the following: (i) the introduction of the interaction between cork growth cycle stage and understory management practices, only possible with cork sample collections from different cork rotation cycles; (ii) the finding that there was no significant effect of intraspecific competition on cork growth